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软骨鱼纲

软骨鱼纲
属 种 发现地点 发现部位 备注 图片
无尖齿鱼科未定属
Acrodontidae indet.[1] 		未定种 属于弓鲛目
Bahariyodon[1] B. bartheli 属于弓鲛目
Cenocarcharias[1] C. tenuiplicatus 一颗牙齿[1] 属于白垩刺甲鲨科
Distobatus[1] D. nutiae 属于弓鲛目
哈氏鲨科（英语：Haimirichia）
Haimirichia[1] 		H. amonensis 一颗牙齿[1] 属于鼠鲨目
Marckgrafia[1] M. lybica 13颗牙齿[1] 属于鳐总目
帆锯鳐属
Onchopristis 		努米底亚帆锯鳐
O. numida[2] 		目前已知体型最大的锯鳐，属于鳐目[3]
帆锯鳐的吻突与牙齿化石
Peyeria[1] P. libyca 3颗牙齿[1] 可能为努米底亚帆锯鳐的次异名
外侧棘鲨属（英语：Tribodus）
Tribodus[1] 		T. sp. 属于弓鲛目

条鳍鱼总纲

条鳍鱼总纲
属 种 发现地点 发现部位 备注 图片
阿德里亚娜鱼属
Adrianaichthys[1] 		A. pankowskii 鳞片[4]与两具颅骨[5] 属于雀鳝目，原本被认为属于鳞齿鱼属（英语：Lepidotes），但之后被重新分类至独立属[6]
埃达克龙鱼属（英语：Aidachar）复原图
Afrocascudo（英语：Afrocascudo）[7] A. saharaensis 属于新鳍亚纲，可能属于甲鲶科或属于雀鳝目奥巴鱼科的幼年个体[8]
Agassizilia（英语：Agassizilia）[9] A. erfoudina 可能属于坚齿鱼科
阿古尔特鱼属
Agoultichthys[1] 		A. chattertoni 属于条鳍鱼总纲，但详细分类位置不明，可能属于巨半椎鱼科（英语：Macrosemiidae）[10]或Ophiopsiellidae[11]
埃达克龙鱼属（英语：Aidachar）
Aidachar 		A. pankowskii 属于乞丐鱼目
巴奇鱼属
Bartschichthys[1] 		B. sp. 背鳍的的刺状鳍骨[1] 属于腕鳍鱼纲
巴氏巴威多鳍鱼
Bawitius 		cf. B. sp. 鳞片与下颚碎片[4] 属于腕鳍鱼纲
卡拉矛普莱弓鳍鱼属（英语：Calamopleurus）
Calamopleurus[1] 		非洲卡拉矛普莱弓鳍鱼
C. africanus 		部分颅骨[1] 属于弓鳍鱼目
Concavotectum（英语：Concavotectum）[1] C. moroccensis 属于塞纳命运鱼目
齿雀鳝属
Dentilepisosteus[1] 		D. kemkemensis 属于雀鳝目
双颌鲱属（英语：Diplomystus）
Diplomystus[1] 		D. sp. 属于骨鲱群真骨类
Diplospondichthys[1] D. moreaui A member of Actinopterygii of uncertain phylogenetic placement, possibly a teleost
Erfoudichthys[1] E. rosae Isolated skull[1] A small-bodied teleost of unknown affinity
Neoproscinetes[9] N. africanus A member of the family Pycnodontidae
Obaichthys O. africanus Isolated scales[4] A member of Lepisosteiformes
Oniichthys O. falipoui Near complete skeleton including skull[4] A member of Lepisosteiformes
Palaeonotopterus[1] P. greenwoodi A member of Osteoglossomorpha
Serenoichthys[1] S. kemkemensis Several articulated skeletons[1] A small cladistian
Spinocaudichthys[1] S. oumtkoutensis An elongate freshwater acanthomorph
Stromerichthys S. aethiopicus
Sudania[1] S. sp. An isolated pinnula[1] A cladistian

肉鳍鱼总纲

肉鳍鱼总纲
属 种 发现地点 发现部位 备注 图片
Arganodus A. tiguidiensis A lungfish
Life restoration of Axelrodichthys
The Queensland Lungfish, the only living member of Neoceratodus
Axelrodichthys[12] A.? lavocati A mawsoniid coelacanth; this species was previously assigned to Mawsonia, and its generic assignment is still not certain[13]
Neoceratodus N. africanus A lungfish

Amphibians

Amphibians
属 种 发现地点 发现部位 备注 图片
Anura indet.[14] Indeterminate Douira Formation Incomplete left ilium
Cretadhefdaa[14] C. taouzensis Douira Formation Posterior portion of the skull, incomplete squamosal, incomplete maxilla, three incomplete presacral vertebrae, one incomplete sacral vertebra A neobatrachian frog with possible hyloid affinities.
cf. Kababisha[15] Indeterminate A salamander belonging to the family Sirenidae
?Neobatrachia indet.[14] Indeterminate Douira Formation Incomplete humerus A frog, possibly a member of Ranoidea.
Oumtkoutia[15] O. anae A frog belonging to the family Pipidae

Lizards and snakes

色键
分类单元 重新分类的分类单元 错误的分类单元 可疑的分类单元或初级同义词 足迹分类单元 蛋分类单元 形态分类群
注释
不确定或暂定的分类单元以小字体展示； 被划掉的分类单元不可信
Lizards and snakes reported from the Continental Red Beds
属 种 发现地点 发现部位 备注 图片

Bicuspidon

B. hogreli[16]

A polyglyphanodontid lizard.

Jeddaherdan[17]

J. aleadonta

Partial mandible with teeth.

An iguanian belonging to the group Acrodonta, possibly a relative of the uromastycine agamids. Argued by Vullo et al. (2022) to actually come from Quaternary beds, and to be based on a fossil material of a member of the genus Uromastyx.[18]

Lapparentophis

L. ragei[19]

Two isolated trunk vertebrae

An early snake.

Madtsoiidae indet.[15]

Indeterminate

Vertebrae[1]

An early snake.

?Nigerophiidae indet.[15]

Indeterminate

Dorsal vertebrae[1]

An early snake.

Norisophis[20]

N. begaa[20]

One posterior and two mid-trunk vertebrae

A stem-snake.

Indeterminate[20]

A mid-trunk vertebra

Simoliophis[15]

cf. S. libycus

Vertebrae[1]

An early snake.

Plesiosaurs

Plesiosaurs
属 种 发现地点 发现部位 备注 图片
Leptocleididae cf. Leptocleidus[21] indeterminate

Turtles

Turtles reported from the Continental Red Beds
属 种 发现地点 发现部位 备注 图片

Dirqadim

D. schaefferi

A Euraxemydid

Galianemys

G. emringeri

A Cearachelyin

G. whitei

Hamadachelys

H. escuilliei

Crocodylomorphs

A tooth enamel identified as cf. Sarcosuchus was discovered from the Ifezouane Formation.[22]

Crocodylomorphs reported from the Continental Red Beds
属 种 发现地点 发现部位 备注 图片

Aegisuchus

A. witmeri

"Partial braincase of a large individual with skull roof, temporal, and occipital regions."[23]

An aegyptosuchid that may be a synonym of Laganosuchus.[1]

Aegisuchus
Araripesuchus
Elosuchus cherifiensis
Laganosuchus

Antaeusuchus[24]

A. taouzensis

Paired mandibles and a partial right mandible

A peirosaurid.

Araripesuchus

A. rattoides

Douira Formation

Elosuchus

E. cherifiensis

  • Gara Sbaa Formation
  • Douira Formation

An elosuchid. The material may represent two different species.[1]

Hamadasuchus

H. rebouli

  • Gara Sbaa Formation
  • Douira Formation?

A peirosaurid.

Kemkemia

K. auditorei

Errachidia Province, Morocco[25]

Known from an isolated caudal vertebra.[25]

Initially thought to be a neotheropod,[25] but subsequently discovered to be an indeterminate crocodyliform.[26]

Laganosuchus

L. thaumastos

A stomatosuchid.

Lavocatchampsa[27]

Lavocatchampsa sigogneaurusselae

Anterior portion of a rostrum with mandible, with an almost complete dentition[27]

A candidodontid notosuchian.[27]

Dinosaurs

Indeterminate lithostrotian remains once misattributed to the Titanosauridae are present in the province of Ksar-es-Souk, Morocco.[28]

色键
分类单元 重新分类的分类单元 错误的分类单元 可疑的分类单元或初级同义词 足迹分类单元 蛋分类单元 形态分类群
注释
不确定或暂定的分类单元以小字体展示； 被划掉的分类单元不可信

Ornithischians

Ornithischians reported from the Continental Red Beds
属 种 发现地点 发现部位 备注 图片

Ankylosauria[1][29]

Indeterminate

Douira Formation

An isolated tooth.[1]

A probable ankylosaur[29]

Ornithopoda[30]

Indeterminate

Douira Formation

A large, clover-shaped, three-toed footprint.[1]

Comparable in size and shape to tracks typically attributed to Iguanodon.[30]

Sauropods

Sauropods reported from the Continental Red Beds
属 种 发现地点 发现部位 备注 图片

Rebbachisaurus

R. garasbae

Ksar-es-Souk province, Morocco.[28]

Gara Sbaa Formation

A rebbachisaurid.

Somphospondyli[31]

Indeterminate

Anterior dorsal vertebra, partial right ischium[31]

The vertebra might belong to a basal titanosaurian, possibly distinct from Aegyptosaurus and Paralititan.[31] The ischium is not identifiable beyond Somphospondyli; it preserves numerous grooves and pits which might be feeding traces left by a very large non-avian theropod.[31]

Titanosauria[32][1]

Indeterminate

  • Ifezouane Formation
  • Douira Formation

Isolated teeth, caudal vertebrae, a partial humerus, a tarsal bone and the proximal end of an ulna.[1]

Fossil material of one or more titanosaurian sauropods. Some fossils are indicative of large body size comparable to Paralititan stromeri.[1]

Theropods

Theropods reported from the Continental Red Beds
属 种 发现地点 发现部位 备注 图片

Abelisauridae[33]

Indeterminate

Isolated teeth.[33][34]
Partial right femur.[35] Partial right maxilla.[1] A cervical vertebra.[36] Left maxilla.[1]

Abelisaurid material belonging to one or two distinct taxa.[1]

Carcharodontosaurus saharicus
Deltadromeus agilis
Rebbachisaurus garasbae
Rugops primus
Spinosaurus aegyptiacus reconstruction

Carcharodontosaurus[28]

C. saharicus[28]

Ksar-es-Souk province, Morocco.[28] Douira Formation Partial skull, including braincase, nasals, postorbitals, jugals, left lacrimal and right maxilla with most teeth.[37]

A carcharodontosaurid theropod.

Carcharodontosauridae[38] Indeterminate Southeast of Taouz, Errachidia Province Ifezouane Formation partial maxilla and partial jugal A carcharodontosaurid theropod different from C. saharicus

Deltadromeus

D. agilis

Gara Sbaa Formation

"Partial skeleton, isolated limb elements."[39]

A noasaurid ceratosaurian or possible neovenatorid carnosaur. May be synonymous with Bahariasaurus.

Dromaeosauridae[33]

Indeterminate

Isolated teeth.[33]

Originally described as teeth of indeterminate dromaeosaurids. Hendrickx et al. (2024) reinterpreted this fossil material as teeth of abelisauroid theropods, including noasaurids and juvenile abelisaurids.[34]

cf. Elaphrosaurus

Indeterminate

Ksar-es-Souk province, Morocco.[28]

Fossils previously referred to cf. Elaphrosaurus are actually indeterminate theropod remains.

Noasauridae[36]

Indeterminate

An anterior cervical vertebra[36]
Isolated teeth[34]

"Osteoporosia"[40] "O. gigantea"[40] A tooth and a possible neural arch from another specimen.[40] A theropod, possibly synonymous with Sauroniops.[41]

Saurischia[42]

Indeterminate

An isolated cervical vertebra.[42]

An indeterminate saurischian.

Sauroniops[43]

S. pachytholus

Ifezouane Formation

"An isolated and almost complete left frontal,[44] and a possible tooth and neural arch from two other specimens."[41]

A carcharodontosaurid distinct from Carcharodontosaurus.[43][44]

Spinosaurus

S. aegyptiacus

Ksar-es-Souk province, Morocco.[28] Douira Formation Partial skeleton, including parts of the skull, neck, torso, and most of the tail and hind limbs. [45]

Numerous isolated bones.

Pterosaurs

Pterosaurs of the Kem Kem Beds
属 种 发现地点 发现部位 备注 图片

Afrotapejara[46]

A. zouhri[46]

Takmout Ifezouane Formation

A fragment of bone interpreted as a fragment of anterior mandibular symphysis,[47] and additional jaw fragments that pertain to the rostrum as well as indeterminate jaw fragments.[48]

A tapejarid pterosaur. Originally believed to belong to either the family Thalassodromidae[49] or an additional specimen of Alanqa saharica.[50]

Afrotapejara zouhri
Akharhynchus martilli
Alanqa saharica
Anhanguera
Coloborhynchus
Leptostomia
Ornithocheirus
Akharhynchus[51] A. martilli Tafilalt Ifezouane Formation A fragment of the anterior part of the premaxillae A tropeognathine anhanguerian.

Alanqa[49]

A. saharica[49]

Ifezouane Formation The holotype is a mandibular symphysis, of different parts of the jaw A pterosaur of uncertain phylogenetic placement, probably an azhdarchid.[48]

Anhanguera[52]

A. cf. piscator[52]

upper Ifezouane Formation

Partial mandibular symphysis[52]

Apatorhamphus[53]

A. gyrostega[53]

Ifezouane Formation

Partial rostrum and mandible, with additional referred jaw fragments[48]

A possible chaoyangopterid azhdarchoid pterosaur.[53] Originally believed to be a possible pteranodontid,[49] a possible dsungaripterid,[54] a possible non-azhdarchid azhdarchoid or nyctosaurid,[54] or a specimen of Alanqa saharica.[50]

Azhdarchidae indet.[54]

Indeterminate[54]

Three middle cervical vertebrae.[54][49]

Averianov (2014) considered these vertebrae to pertain to Alanqa saharica,[50] although the vertebrae may be indicative of two taxa.[54]

Coloborhynchus[52] C. sp. A.[52] Hassi El Begaa Premaxillae fragment[52] Possibly a specimen of Nicorhynchus fluviferox.[55]

Leptostomia[56]

L. begaaensis[56]

Aferdou N’ Chaft

upper Ifezouane Formation

Partial rostrum and partial mandibular synthesis[56]

A small, long-beaked pterosaur, likely a member of Azhdarchoidea.[56]

Nicorhynchus[55]

N. fluviferox[57][55]

Possibly Aferdou N’Chaft, Hassi El Begaa[55]

Ifezouane Formation

An anterior portion of the rostrum.[55]

Originally described as a species of Coloborhynchus[57] but subsequently transferred to the genus Nicorhynchus.

Ornithocheirus[52]

O. cf. simus.[52]

upper Ifezouane Formation

Premaxillae fragment[52]

Siroccopteryx[58]

S. moroccensis[58]

Anterior part of a rostrum

Classified by some authors as a species belonging to the genus Coloborhynchus.[49]

Xericeps X. curvirostra Aferdou N'Chaft Ifezouane Formation Mandibular symphysis and partial mandible[48] An indeterminate azhdarchoid, possibly a chaoyangopterid.[48]
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